Too loose and they run all day. Too rough and you either have a roughed up dog with injuries, a busted bay or both. How much is genetic.
Vs. learned?
I would have to go with mostly genetic. But an old gritty/rough dog can learn to wear a hog down and be cautious doing it under normal circumstances and then increase the pressure when the hog has slowed down. That would be learned behavior...but then, if the dog learned that on its own then it probably is genetic to a certain extent. It is almost like what came first? The chicken or the egg...